About the Authors

Michael W. Miller

mike.miller@state.co.us

Affiliation Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado, United States of America

Heather M. Swanson

Affiliation City of Boulder Open Space and Mountain Parks, Boulder, Colorado, United States of America

Lisa L. Wolfe

Affiliation Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado, United States of America

Fred G. Quartarone

Affiliation Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado, United States of America

Sherri L. Huwer

Affiliation Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado, United States of America

Charles H. Southwick

Affiliation Department of Ecology and Evolutionary Biology, University of Colorado at Boulder, Boulder, Colorado, United States of America

Paul M. Lukacs

Affiliation Colorado Division of Wildlife, Wildlife Research Center, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MWM HMS LLW SLH CHS PML. Performed the experiments: MWM HMS LLW FGQ. Analyzed the data: MWM PML. Wrote the paper: MWM HMS LLW FGQ SLH CHS PML.