About the Authors

Lu Xu

Contributed equally to this work with: Lu Xu, Hao Wang, Qiong La

Affiliations Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China, College of Life Sciences, Northwest Normal University, Lanzhou, China

Hao Wang

Contributed equally to this work with: Lu Xu, Hao Wang, Qiong La

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Qiong La

Contributed equally to this work with: Lu Xu, Hao Wang, Qiong La

Affiliations Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China, Department of Biology, Tibet University, Lhasa, China

Fan Lu

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Kun Sun

Affiliation College of Life Sciences, Northwest Normal University, Lanzhou, China

Yang Fang

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Mei Yang

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Yang Zhong

Affiliations Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China, Department of Biology, Tibet University, Lhasa, China

Qianhong Wu

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Jiakuan Chen

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

H. John B. Birks

Affiliations Department of Biology, University of Bergen, Bergen, Norway, Environmental Change Research Centre, University College London, London, United Kingdom, School of Geography and the Environment, University of Oxford, Oxford, United Kingdom

Wenju Zhang

wjzhang@fudan.edu.cn

Affiliation Institute of Biodiversity Science, School of Life Sciences, Fudan University, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WZ. Performed the experiments: WZ QL FL HW YZ YF LX. Analyzed the data: MY LX HW WZ. Contributed reagents/materials/analysis tools: LX HW MY YZ KS JC QW. Wrote the paper: WZ LX HW HJBB. Obtained materials: WZ KS HW FL.