About the Authors

Georgi Hudjashov

hudja@ebc.ee

Affiliations Evolutionary Biology Group, Estonian Biocentre, Tartu, Estonia, Department of Evolutionary Biology, Institute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia

Richard Villems

Affiliations Evolutionary Biology Group, Estonian Biocentre, Tartu, Estonia, Department of Evolutionary Biology, Institute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia, Estonian Academy of Sciences, Tallinn, Estonia

Toomas Kivisild

Affiliations Evolutionary Biology Group, Estonian Biocentre, Tartu, Estonia, Department of Evolutionary Biology, Institute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia, Division of Biological Anthropology, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors confirm that Toomas Kivisild is a PLOS ONE Editorial Board member. This does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GH RV TK. Performed the experiments: GH. Analyzed the data: GH TK. Contributed reagents/materials/analysis tools: TK RV. Wrote the manuscript: GH TK RV.