About the Authors

Yuk-Sang Chan

Contributed equally to this work with: Yuk-Sang Chan, Ryo Takeuchi, Jordan Jarjour, David S. Huen

Affiliation Department of Genetics, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Ryo Takeuchi

Contributed equally to this work with: Yuk-Sang Chan, Ryo Takeuchi, Jordan Jarjour, David S. Huen

Affiliations Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Northwest Genome Engineering Consortium, Seattle, Washington, United States of America

Jordan Jarjour

Contributed equally to this work with: Yuk-Sang Chan, Ryo Takeuchi, Jordan Jarjour, David S. Huen

Affiliation Pregenen Inc., Seattle, Washington, United States of America

David S. Huen

Contributed equally to this work with: Yuk-Sang Chan, Ryo Takeuchi, Jordan Jarjour, David S. Huen

david.s.huen@gmail.com

Current address: Department of Applied Sciences, University of Wolverhampton, Wolverhampton, United Kingdom

Affiliation Department of Genetics, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Barry L. Stoddard

Affiliations Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Northwest Genome Engineering Consortium, Seattle, Washington, United States of America

Steven Russell

Affiliations Department of Genetics, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom, Cambridge Systems Biology Centre, Cambridge, Cambridgeshire, United Kingdom

Competing Interests

The authors have read the journal's policy and have the following potential conflicts: BLS is a founder, and JJ an employee and shareholder, of a biotechnology startup company (Pregenen Inc) that manufactures homing endonuclease variants of I-OnuI as described in this manuscript. Neither individual nor that company have any commercial stake or interest in the constructs described in this manuscript. The authors do not have patents on the material within the paper. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YC RT JJ DSH BLS SR. Performed the experiments: YC RT JJ DSH. Analyzed the data: YC RT JJ DSH BLS SR. Contributed reagents/materials/analysis tools: YC RT JJ DSH. Wrote the paper: RT JJ DSH BLS SR.