About the Authors

Stephanie N. Lewis

Affiliations Genetics, Bioinformatics, and Computational Biology Program, Virginia Tech, Blacksburg, Virginia, United States of America, Department of Biochemistry, Virginia Tech, Blacksburg, Virginia, United States of America

Elaine Nsoesie

Affiliation Genetics, Bioinformatics, and Computational Biology Program, Virginia Tech, Blacksburg, Virginia, United States of America

Charles Weeks

Affiliation Genetics, Bioinformatics, and Computational Biology Program, Virginia Tech, Blacksburg, Virginia, United States of America

Dan Qiao

Affiliation Department of Computer Science, Virginia Tech, Blacksburg, Virginia, United States of America

Liqing Zhang

lzhang@systemsbiology.org

Affiliations Genetics, Bioinformatics, and Computational Biology Program, Virginia Tech, Blacksburg, Virginia, United States of America, Department of Computer Science, Virginia Tech, Blacksburg, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LZ SNL EN DQ CW. Performed the experiments: SNL EN DQ CW. Analyzed the data: SNL EN DQ CW LZ. Wrote the paper: SNL EN DQ CW. Revised the manuscript: LZ SNL EN.