About the Authors

Mayumi Oakland

Current address: Department of Microbiology, University of Iowa, Iowa City, Iowa, United States of America

Affiliation Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, United States of America

Byeonghwa Jeon

Current address: Department of Pathology and Microbiology, Atlantic Veterinary College, University of Prince Edward Island, Charlottetown, Prince Edward Island, Canada

Affiliation Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, United States of America

Orhan Sahin

Affiliation Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, United States of America

Zhangqi Shen

Affiliation Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, United States of America

Qijing Zhang

zhang123@iastate.edu

Affiliation Department of Veterinary Microbiology and Preventive Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MO BJ OS QZ. Performed the experiments: MO BJ OS ZS. Analyzed the data: MO BJ OS ZS QZ. Contributed reagents/materials/analysis tools: MO BJ OS ZS QZ. Wrote the paper: MO QZ.