About the Authors

Richard Stafford

rstafford@glos.ac.uk

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Adam G. Hart

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Laura Collins

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Claire L. Kirkhope

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Rachel L. Williams

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Samuel G. Rees

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Jane R. Lloyd

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Anne E. Goodenough

Affiliation Department of Natural and Social Sciences, University of Gloucestershire, Cheltenham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS AH JL AG. Performed the experiments: LC CK RW SR. Analyzed the data: CK RW SR. Contributed reagents/materials/analysis tools: RS JL. Wrote the paper: RS AH LC AG.