About the Authors

Yong Zhao

Contributed equally to this work with: Yong Zhao, Dongsheng Xu

Affiliation School of Biological Sciences, University of Nebraska, Lincoln, Nebraska, United States of America

Dongsheng Xu

Contributed equally to this work with: Yong Zhao, Dongsheng Xu

Affiliation School of Biological Sciences, University of Nebraska, Lincoln, Nebraska, United States of America

Yanjun Jiang

Affiliation School of Biological Sciences, University of Nebraska, Lincoln, Nebraska, United States of America

Luwen Zhang

lzhang2@unlnotes.unl.edu

Affiliations School of Biological Sciences, University of Nebraska, Lincoln, Nebraska, United States of America, Nebraska Center for Virology, University of Nebraska, Lincoln, Nebraska, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YZ DX LZ. Performed the experiments: YZ DX YJ. Analyzed the data: YZ DX LZ. Contributed reagents/materials/analysis tools: YZ DX YJ LZ. Wrote the paper: LZ.