About the Authors

N Schweighofer

To whom correspondence should be addressed. E-mail: schweigh@usc.edu

Affiliation Biokinesiology and Physical Therapy, University of Southern California, Los Angeles, United States of America

K Shishida

Affiliation Department of Psychiatry and Neurosciences, Hiroshima University, Hiroshima, Japan

C. E Han

Affiliation Computer Science, University of Southern California, Los Angeles, United States of America

Y Okamoto

Affiliation Department of Psychiatry and Neurosciences, Hiroshima University, Hiroshima, Japan

S. C Tanaka

Affiliations Bioinformatics and Genomics, Nara Institute of Science and Technology, Nara, Japan , Department of Computational Neurobiology, Advanced Telecommunications Research Institute International Computational Neuroscience Labs, Kyoto, Japan

S Yamawaki

Affiliation Department of Psychiatry and Neurosciences, Hiroshima University, Hiroshima, Japan

K Doya

Affiliations Department of Computational Neurobiology, Advanced Telecommunications Research Institute International Computational Neuroscience Labs, Kyoto, Japan , Initial Research Project, Okinawa Institute of Science and Technology, Okinawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

NS, KS, YO, SCT, SY, and KD conceived and designed the experiments. KS, YO, and SCT performed the experiments. NS and CEH analyzed the data. NS wrote the paper.