About the Authors

Yusuke Nakano

Roles Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – review & editing

Affiliation Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan

Keisuke Yamamoto

Roles Investigation, Methodology, Validation, Visualization, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Mahoko Takahashi Ueda

Roles Data curation, Formal analysis, Investigation, Software, Visualization, Writing – review & editing

Affiliation Department of Molecular Life Science, Tokai University School of Medicine, Kanagawa, Japan

Andrew Soper

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yoriyuki Konno

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Izumi Kimura

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Keiya Uriu

Roles Investigation, Writing – review & editing

Affiliations Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan

Ryuichi Kumata

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Faculty of Science, Kyoto University, Kyoto, Japan

Hirofumi Aso

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Naoko Misawa

Roles Investigation, Methodology, Visualization, Writing – review & editing

Affiliation Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan

Shumpei Nagaoka

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Soma Shimizu

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Keito Mitsumune

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Yusuke Kosugi

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Guillermo Juarez-Fernandez

Roles Investigation, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Jumpei Ito

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan

So Nakagawa

Roles Data curation, Formal analysis, Funding acquisition, Resources, Software, Writing – review & editing

Affiliation Department of Molecular Life Science, Tokai University School of Medicine, Kanagawa, Japan

Terumasa Ikeda

Roles Funding acquisition, Investigation, Resources, Writing – review & editing

Affiliations Department of Biochemistry, Molecular Biology and Biophysics, Masonic Cancer Center, Institute for Molecular Virology, Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, University of Minnesota, Minneapolis, Minnesota, United States of America, Division of Molecular Virology and Genetics, Joint Research Center for Human Retrovirus Infection, Kumamoto University, Kumamoto, Japan

Yoshio Koyanagi

Roles Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Medicine, Kyoto University, Kyoto, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan

Reuben S. Harris

Roles Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliations Department of Biochemistry, Molecular Biology and Biophysics, Masonic Cancer Center, Institute for Molecular Virology, Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America, Howard Hughes Medical Institute, University of Minnesota, Minneapolis, Minnesota, United States of America

Kei Sato

Roles Conceptualization, Funding acquisition, Project administration, Supervision, Visualization, Writing – original draft, Writing – review & editing

ksato@ims.u-tokyo.ac.jp

Affiliations Laboratory of Systems Virology, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto, Japan, Division of Systems Virology, Department of Infectious Disease Control, International Research Center for Infectious Diseases, Institute of Medical Science, the University of Tokyo, Tokyo, Japan, Graduate School of Pharmaceutical Sciences, Kyoto University, Kyoto, Japan, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan, CREST, Japan Science and Technology Agency, Saitama, Japan

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: R.S.H. is a co-founder, shareholder and consultant of ApoGen Biotechnologies.