About the Authors

Benjamin R. Treat

Contributed equally to this work with: Benjamin R. Treat, Sarah M. Bidula

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Molecular Virology and Microbiology Graduate Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Sarah M. Bidula

Contributed equally to this work with: Benjamin R. Treat, Sarah M. Bidula

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft

Affiliations Molecular Virology and Microbiology Graduate Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Srividya Ramachandran

Roles Investigation, Methodology

Current address: Neurological Institute, Cleveland Clinic, Cleveland, Ohio, United States of America

Affiliations Molecular Virology and Microbiology Graduate Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Anthony J. St Leger

Roles Investigation, Methodology

Current address: Immunoregulation Section, NEI, NIH, Bethesda, Maryland, United States of America

Affiliations Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Immunology Graduate Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Robert L. Hendricks

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft, Writing – review & editing

Affiliations Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Immunology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, Pittsburgh, Pennsylvania, United States of America

Paul R. Kinchington

Roles Conceptualization, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

Kinchingtonp@upmc.edu

Affiliations Molecular Virology and Microbiology Graduate Program, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Ophthalmology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.