About the Authors

Takasuke Fukuhara

Contributed equally to this work with: Takasuke Fukuhara, Tomokazu Tamura

Roles Conceptualization, Formal analysis, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Tomokazu Tamura

Contributed equally to this work with: Takasuke Fukuhara, Tomokazu Tamura

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft

Affiliations Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan, Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Hokkaido, Japan

Chikako Ono

Roles Investigation

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Mai Shiokawa

Roles Investigation

Affiliation School of Veterinary Nursing and Technology, Faculty of Veterinary Science, Nippon Veterinary and Life Science University, Tokyo, Japan

Hiroyuki Mori

Roles Investigation

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Kentaro Uemura

Roles Investigation

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Satomi Yamamoto

Roles Investigation, Resources

Current address: Laboratory of Veterinary Microbiology, School of Veterinary Medicine, Kitasato University, Aomori, Japan

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Takeshi Kurihara

Roles Investigation

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Toru Okamoto

Roles Investigation

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Ryosuke Suzuki

Roles Methodology, Resources

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Kentaro Yoshii

Roles Resources, Supervision

Affiliation Laboratory of Public Health, Department of Environmental Veterinary Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Hokkaido, Japan

Takeshi Kurosu

Roles Resources

Affiliation Department of Virology I, National Institute of Infectious Diseases, Tokyo, Japan

Manabu Igarashi

Roles Investigation, Supervision

Affiliations Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Hokkaido, Japan, Division of Global Epidemiology, Research Center for Zoonosis Control, Hokkaido University, Hokkaido, Japan

Hiroshi Aoki

Roles Conceptualization, Resources

Affiliation School of Veterinary Nursing and Technology, Faculty of Veterinary Science, Nippon Veterinary and Life Science University, Tokyo, Japan

Yoshihiro Sakoda

Roles Conceptualization

Affiliations Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Hokkaido, Japan, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Hokkaido, Japan

Yoshiharu Matsuura

Roles Conceptualization, Funding acquisition, Resources, Writing – original draft, Writing – review & editing

matsuura@biken.osaka-u.ac.jp

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Competing Interests

The authors have declared that no competing interest exist.