About the Authors

Mary Jane Drake

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Benjamin Brennan

Affiliation MRC-University of Glasgow Centre for Virus Research, University of Glasgow, Glasgow, United Kingdom

Kenneth Briley Jr

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Stephen M. Bart

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Eric Sherman

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Agnieszka M. Szemiel

Affiliation MRC-University of Glasgow Centre for Virus Research, University of Glasgow, Glasgow, United Kingdom

Madeleine Minutillo

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Frederic D. Bushman

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Paul Bates

pbates@upenn.edu

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: MJD KB PB. Formal analysis: MJD BB SMB ES FDB PB. Funding acquisition: FDB PB. Investigation: MJD BB KB SMB ES AMS MM. Methodology: MJD BB ES FDB PB. Project administration: MJD PB. Resources: MJD BB KB ES FDB PB. Software: ES FDB. Supervision: PB. Visualization: MJD. Writing – original draft: MJD PB. Writing – review & editing: MJD BB KB FDB PB.