About the Authors

Nathan D. Grubaugh

Contributed equally to this work with: Nathan D. Grubaugh, Darci R. Smith

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Darci R. Smith

Contributed equally to this work with: Nathan D. Grubaugh, Darci R. Smith

Current address: Southern Research Institute, Frederick, Maryland, United States of America

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Doug E. Brackney

¤b Current address: The Connecticut Agricultural Experiment Station, New Haven, Connecticut, United States of America

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Angela M. Bosco-Lauth

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Joseph R. Fauver

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Corey L. Campbell

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Todd A. Felix

Affiliation United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, Lakewood, Colorado, United States of America

Hannah Romo

Affiliations Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America, Division of Vector-Borne Diseases, Centers for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Nisha K. Duggal

Affiliation Division of Vector-Borne Diseases, Centers for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Elizabeth A. Dietrich

Affiliation Division of Vector-Borne Diseases, Centers for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Tyler Eike

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Jennifer E. Beane

Affiliation Section for Computational Biomedicine, Boston University School of Medicine, Boston, Massachusetts, United States of America

Richard A. Bowen

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

William C. Black

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Aaron C. Brault

Affiliation Division of Vector-Borne Diseases, Centers for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Gregory D. Ebel

gregory.ebel@colostate.edu

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GDE DRS NDG. Performed the experiments: DRS NDG ACB DEB AMBL JRF TAF CLC HR NKD EAD RAB. Analyzed the data: NDG GDE WCB DRS JEB TE. Contributed reagents/materials/analysis tools: RAB TAF ACB JEB GDE. Wrote the paper: NDG GDE DRS.