About the Authors

Dipu Mohan Kumar

Affiliation Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, Ohio, United States of America

Mamoru Yamaguchi

Affiliation Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, Ohio, United States of America

Koshiro Miura

Current address: Department of Nutrition, Kyushu Women's University, Kita-kyushu City, Fukuoka, Japan

Affiliation Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, Ohio, United States of America

Mingqun Lin

Affiliation Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, Ohio, United States of America

Marek Los

Affiliation Department of Clinical & Experimental Medicine, Integrative Regenerative Medical Center Linköping University, Linkoping, Sweden

Johannes F. Coy

Affiliation Vorstand/CSO TAVARLIN AG, Darmstadt, Germany

Yasuko Rikihisa

rikihisa.1@osu.edu

Affiliation Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State University, Columbus, Ohio, United States of America

Competing Interests

JFC is employed by Vorstand/CSO TAVARLIN AG and has patent applications (pending), including individual applications or those belonging to the institution to which he is affiliated. This does not alter our adherence to all PLOS Pathogens policies on sharing data and materials. All other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YR DMK. Performed the experiments: DMK YR MY KM. Analyzed the data: DMK YR MLi. Contributed reagents/materials/analysis tools: MLo JFC. Wrote the paper: YR DMK.