About the Authors

Kristin L. Shingler

Affiliation Department of Medicine, Department of Microbiology and Immunology, The Pennsylvania State University College of Medicine, Hershey, Pennsylvania, United States of America

Jennifer L. Yoder

Affiliation Department of Medicine, Department of Microbiology and Immunology, The Pennsylvania State University College of Medicine, Hershey, Pennsylvania, United States of America

Michael S. Carnegie

Affiliation Department of Medicine, Department of Microbiology and Immunology, The Pennsylvania State University College of Medicine, Hershey, Pennsylvania, United States of America

Robert E. Ashley

Affiliation Department of Medicine, Department of Microbiology and Immunology, The Pennsylvania State University College of Medicine, Hershey, Pennsylvania, United States of America

Alexander M. Makhov

Affiliation Department of Structural Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

James F. Conway

Affiliation Department of Structural Biology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Susan Hafenstein

shafenstein@hmc.psu.edu

Affiliation Department of Medicine, Department of Microbiology and Immunology, The Pennsylvania State University College of Medicine, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KLS SH. Performed the experiments: KLS. Analyzed the data: KLS SH. Contributed reagents/materials/analysis tools: JLY MSC REA AMM JFC. Wrote the paper: KLS SH.