About the Authors

Wen-Ping Guo

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Xian-Dan Lin

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation Wenzhou Center for Disease Control and Prevention, Wenzhou, Zhejiang Province, China

Wen Wang

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Jun-Hua Tian

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation Wuhan Center for Disease Control and Prevention, Wuhan, Hubei Province, China

Mei-Li Cong

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Hai-Lin Zhang

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation Yunnan Institute of Endemic Diseases Control and Prevention, Dali, China

Miao-Ruo Wang

Contributed equally to this work with: Wen-Ping Guo, Xian-Dan Lin, Wen Wang, Jun-Hua Tian, Mei-Li Cong, Hai-Lin Zhang, Miao-Ruo Wang

Affiliation Longquan Center for Disease Control and Prevention, Longquan, Zhejiang Province, China

Run-Hong Zhou

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Jian-Bo Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Ming-Hui Li

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Jianguo Xu

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Edward C. Holmes

Affiliations Sydney Emerging Infections and Biosecurity Institute, School of Biological Sciences and Sydney Medical School, The University of Sydney, Sydney, Australia, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Yong-Zhen Zhang

zhangyongzhen@icdc.cn

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Department of Zoonoses, National Institute for Communicable Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Changping, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YZZ. Performed the experiments: WPG WW MLC RHZ MHL XDL JHT HLZ MRW JBW. Analyzed the data: YZZ WPG WW RHZ JX. Wrote the paper: YZZ ECH. Collected animal samples: XDL JHT HLZ MRW JBW.