About the Authors

Justin Lessler

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Steven Riley

s.riley@imperial.ac.uk

Affiliation School of Public Health, Department of Infectious Disease Epidemiology, MRC Centre for Outbreak Analysis and Modelling, Imperial College, London, United Kingdom

Jonathan M. Read

Affiliation Department of Epidemiology and Population Health, Institute of Infection and Global Health, Faculty of Health and Life Sciences, University of Liverpool, Liverpool, United Kingdom

Shuying Wang

Affiliation Guangzhou No. 12 Hospital, Guangzhou, Guangdong, China

Huachen Zhu

Affiliations International Institute of Infection and Immunity, Shantou University Medical College, Shantou, Guangdong, China, Department of Microbiology, The University of Hong Kong, Hong Kong SAR, China

Gavin J. D. Smith

Affiliation Laboratory of Virus Evolution, Program in Emerging Infectious Diseases, Duke-NUS Graduate Medical School, Singapore, Singapore

Yi Guan

Affiliations International Institute of Infection and Immunity, Shantou University Medical College, Shantou, Guangdong, China, Department of Microbiology, The University of Hong Kong, Hong Kong SAR, China

Chao Qiang Jiang

Affiliation Guangzhou No. 12 Hospital, Guangzhou, Guangdong, China

Derek A. T. Cummings

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SR DATC JMR GJDS CQJ YG. Performed the experiments: SW HZ. Analyzed the data: SR DATC JMR. Wrote the paper: JL SR JMR DATC.