About the Authors

G. Adam Mott

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Raymond Wilson

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Anuruddika Fernando

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Ailie Robinson

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Paula MacGregor

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

David Kennedy

Affiliation Moredun Research Institute, Pentlands Science Park, Edinburgh, United Kingdom

Dick Schaap

Affiliation Intervet Schering-Plough, Boxmeer, The Netherlands

Jacqueline B. Matthews

Affiliation Moredun Research Institute, Pentlands Science Park, Edinburgh, United Kingdom

Keith R. Matthews

keith.matthews@ed.ac.uk

Affiliation Centre for Immunity, Infection and Evolution, Institute for Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

Work in this manuscript received contributory (10%) funding from Intervet Schering Plough as part of a contribution to a UK BBSRC Industrial Partnership Award. The information contained within the manuscript is protected by a patent to the University of Edinburgh.

Author Contributions

Conceived and designed the experiments: KRM GAM RW. Performed the experiments: GAM RW AF AR PM DK. Analyzed the data: KRM GAM RW DS JBM. Contributed reagents/materials/analysis tools: DS. Wrote the paper: KRM GAM.