About the Authors

Angela M. Jansen

Contributed equally to this work with: Angela M. Jansen, Lindsay J. Hall

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Lindsay J. Hall

Contributed equally to this work with: Angela M. Jansen, Lindsay J. Hall

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Simon Clare

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

David Goulding

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Kathryn E. Holt

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Andrew J. Grant

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Piero Mastroeni

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Gordon Dougan

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Robert A. Kingsley

rak@sanger.ac.uk

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMJ LJH GD RAK. Performed the experiments: AMJ LJH SC DG. Analyzed the data: AMJ LJH KEH AJG PM GD RAK. Wrote the paper: AMJ LJH RAK.