About the Authors

Martha I. Nelson

nelsonma@mail.nih.gov

Affiliation Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Philippe Lemey

Affiliation Department of Microbiology and Immunology, Katholieke Universiteit Leuven, Leuven, Belgium

Yi Tan

Affiliation Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Amy Vincent

Affiliation Virus and Prion Diseases of Livestock Research Unit, National Animal Disease Center, USDA-ARS, Ames, Iowa, United States of America

Tommy Tsan-Yuk Lam

Affiliation Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Susan Detmer

Affiliation The University of Minnesota Veterinary Diagnostic Laboratory, St. Paul, Minnesota, United States of America

Cécile Viboud

Affiliation Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Marc A. Suchard

Affiliation Departments of Biomathematics and Human Genetics, David Geffen School of Medicine at UCLA, and Department of Biostatistics, UCLA School of Public Health, Los Angeles, California, United States of America

Andrew Rambaut

Affiliations Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America, Institute of Evolutionary Biology, University of Edinburgh, Ashworth Laboratories, Edinburgh, United Kingdom

Edward C. Holmes

Affiliations Division of International Epidemiology and Population Studies, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Marie Gramer

Affiliation The University of Minnesota Veterinary Diagnostic Laboratory, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MIN PL MAS AR ECH. Performed the experiments: AV SD MG. Analyzed the data: MIN PL YT TTYL MAS CV AR ECH. Wrote the paper: MIN PL AV CV MAS AR ECH MG.