About the Authors

J. Joukje Siebenga

Contributed equally to this work with: J. Joukje Siebenga, Philippe Lemey

joukjes@yahoo.co.uk

Affiliations National Institute for Public Health and the Environment, RIVM, Bilthoven, The Netherlands, Erasmus Medical Centre, Rotterdam, The Netherlands

Philippe Lemey

Contributed equally to this work with: J. Joukje Siebenga, Philippe Lemey

Affiliation Rega Institute, Katholieke Universiteit Leuven, Leuven, Belgium

Sergei L. Kosakovsky Pond

Affiliation Antiviral Research Center, University of California San Diego, San Diego, California, United States of America

Andrew Rambaut

Affiliation Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom, and Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Harry Vennema

Affiliation National Institute for Public Health and the Environment, RIVM, Bilthoven, The Netherlands

Marion Koopmans

Affiliations National Institute for Public Health and the Environment, RIVM, Bilthoven, The Netherlands, Erasmus Medical Centre, Rotterdam, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJS PL HV MK. Performed the experiments: JJS PL SLKP. Analyzed the data: JJS PL SLKP HV. Contributed reagents/materials/analysis tools: JJS PL SLKP AR. Wrote the paper: JJS PL.