About the Authors

Matthew D Dyer

Affiliations Genetics, Bioinformatics, and Computational Biology Program, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America , Virginia Bioinformatics Institute, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

T. M Murali

To whom correspondence should be addressed. E-mail: murali@cs.vt.edu (TMM), sobral@vbi.vt.edu (BWS)

Affiliation Department of Computer Science, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Bruno W Sobral

To whom correspondence should be addressed. E-mail: murali@cs.vt.edu (TMM), sobral@vbi.vt.edu (BWS)

Affiliation Virginia Bioinformatics Institute, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TMM proposed the study. MDD and TMM designed the analysis. MDD gathered the data and performed the analysis. MDD, TMM, and BWS analyzed the results and wrote the manuscript.