About the Authors

Meg M Laakso

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Fang-Hua Lee

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Beth Haggarty

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Caroline Agrawal

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Katrina M Nolan

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Mark Biscone

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Josephine Romano

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Andrea P. O Jordan

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

George J Leslie

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Eric G Meissner

Affiliations Department of Microbiology and Immunology, The University of North Carolina, Chapel Hill, North Carolina, United States of America , The Lineberger Comprehensive Cancer Center, The University of North Carolina, Chapel Hill, North Carolina, United States of America

Lishan Su

Affiliations Department of Microbiology and Immunology, The University of North Carolina, Chapel Hill, North Carolina, United States of America , The Lineberger Comprehensive Cancer Center, The University of North Carolina, Chapel Hill, North Carolina, United States of America

James A Hoxie

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Robert W Doms

To whom correspondence should be addressed. E-mail: doms@mail.med.upenn.edu

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MML, FHL, KMN, EGM, JAH, and RWD conceived and designed the experiments. MML, FHL, BH, CA, KMN, JR, and APOJ performed the experiments. MML, FHL, JR, APOJ, JAH, and RWD analyzed the data. FHL, MB, GJL, EGM, and LS contributed reagents/materials/analysis tools. MML and RWD wrote the paper.