About the Authors

Patrick M Ferree

Contributed equally to this work with: Patrick M Ferree, Horacio M Frydman

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of California, Santa Cruz, California, United States of America

Horacio M Frydman

Contributed equally to this work with: Patrick M Ferree, Horacio M Frydman

Affiliation Howard Hughes Medical Institute, Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Jennifer M Li

Affiliation Howard Hughes Medical Institute, Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Jian Cao

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of California, Santa Cruz, California, United States of America

Eric Wieschaus

Affiliation Howard Hughes Medical Institute, Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

William Sullivan

To whom correspondence should be addressed. E-mail: sullivan@biology.ucsc.edu

Affiliation Department of Molecular, Cellular, and Developmental Biology, University of California, Santa Cruz, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PMF, HMF, EW, and WS conceived and designed the experiments. PMF, HMF, JML, and JC performed the experiments. PMF, HMF, JML, JC, EW, and WS analyzed the data. PMF, HMF, EW, and WS contributed reagents/materials/analysis tools. PMF wrote the paper. HMF, EW, and WS edited the paper.