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School of Biological Sciences, Life Sciences Building, University of Bristol, 24 Tyndall Avenue, Bristol, BS8 1TQ, United Kingdom
British Trust for Ornithology, The Nunnery, Thetford, Norfolk, IP24 2PU, United Kingdom
Graduate School of Education, and Centre for Multilevel Modelling, University of Bristol, 2 Priory Road, Bristol, BS8 1TX, United Kingdom
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