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Contributed equally to this work with:
Primate Research Institute, Kyoto University, Inuyama, Japan
Lukuru Wildlife Research Foundation, Kinshasa, Democratic Republic of Congo
Bonobo and Congo Biodiversity Initiative, Zoological Society of Milwaukee, Milwaukee, Wisconsin, United States of America
African Wildlife Foundation, Kinshasa, Democratic Republic of Congo
African Wildlife Foundation, Kinshasa, Democratic Republic of Congo,
Department of Anthropology, Emory University, Atlanta, Georgia, United States of America
Research Center for Ecology and Forestry, Ministry of High Education and Scientific Research, Mabali, Democratic Republic of Congo
World Wide Fund for Nature, Kinshasa, Democratic Republic of Congo
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