About the Authors

Chiyo Kitayama

Roles Conceptualization, Formal analysis, Investigation, Project administration, Writing – original draft

Affiliation Everlasting Nature of Asia (ELNA), Ogasawara Marine Center, Ogasawara, Tokyo, Japan

Keiichi Ueda

Roles Formal analysis, Investigation, Resources, Writing – review & editing

Affiliations Okinawa Churaumi Aquarium, Motobu, Okinawa, Japan, Okinawa Churashima Research Center, Motobu, Okinawa, Japan

Mariko Omata

Roles Investigation, Resources, Writing – review & editing

Affiliation Okinawa Churaumi Aquarium, Motobu, Okinawa, Japan

Taketeru Tomita

Roles Formal analysis, Investigation, Resources, Writing – review & editing

Affiliations Okinawa Churaumi Aquarium, Motobu, Okinawa, Japan, Okinawa Churashima Research Center, Motobu, Okinawa, Japan

Shingo Fukada

Roles Investigation, Resources, Writing – review & editing

Affiliation Okinawa Churaumi Aquarium, Motobu, Okinawa, Japan

Shogo Murakami

Roles Investigation, Resources, Writing – review & editing

Affiliation Shimane Aquarium, Hamada, Shimane, Japan

Yoshiaki Tanaka

Roles Investigation, Resources, Writing – review & editing

Affiliation Shimane Aquarium, Hamada, Shimane, Japan

Akihiro Kaji

Roles Investigation, Resources, Writing – review & editing

Affiliation Shimane Aquarium, Hamada, Shimane, Japan

Satomi Kondo

Roles Writing – review & editing

Affiliation Everlasting Nature of Asia (ELNA), Ogasawara Marine Center, Ogasawara, Tokyo, Japan

Hiroyuki Suganuma

Roles Writing – review & editing

Affiliation Everlasting Nature of Asia (ELNA), Yokohama, Kanagawa, Japan

Yuki Aiko

Roles Formal analysis, Investigation

Affiliation Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Atsuru Fujimoto

Roles Formal analysis, Methodology

Affiliation Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Yusuke K. Kawai

Roles Formal analysis, Methodology

Affiliation Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Masashi Yanagawa

Roles Investigation, Methodology

Affiliation Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Daisuke Kondoh

Roles Conceptualization, Formal analysis, Methodology, Supervision, Writing – original draft, Writing – review & editing

kondoh-d@obihiro.ac.jp

Affiliation Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.