About the Authors

Jennifer L. Essler

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

Affiliation Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Sarah A. Kane

Roles Conceptualization, Data curation, Formal analysis, Methodology, Writing – original draft, Writing – review & editing

Affiliation Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Pat Nolan

Roles Data curation, Project administration, Resources, Supervision, Writing – original draft

Affiliation Tactical Directional Canine, Smithsburg, MD, United States of America

Elikplim H. Akaho

Roles Data curation, Resources, Writing – review & editing

Affiliation Department of Pediatrics, The Children’s Hospital of Philadelphia, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Amalia Z. Berna

Roles Data curation, Resources, Writing – review & editing

Affiliation Department of Pediatrics, The Children’s Hospital of Philadelphia, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Annemarie DeAngelo

Roles Conceptualization, Data curation, Investigation, Methodology, Supervision, Writing – review & editing

Affiliation Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Richard A. Berk

Roles Conceptualization, Formal analysis, Supervision, Writing – review & editing

Affiliations Department of Criminology, School of Arts and Sciences, University of Pennsylvania, Philadelphia, PA, United States of America, Department of Statistics, The Wharton School, University of Pennsylvania, Philadelphia, PA, United States of America

Patricia Kaynaroglu

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Victoria L. Plymouth

Roles Data curation, Formal analysis, Writing – original draft, Writing – review & editing

Affiliation Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Ian D. Frank

Roles Data curation, Resources, Writing – review & editing

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, PA, United States of America

Susan R. Weiss

Roles Data curation, Resources, Writing – review & editing

Affiliation Department of Microbiology, Penn Center for Research on Coronaviruses and Other Emerging Pathogens, University of Pennsylvania School of Medicine, Philadelphia, PA, United States of America

Audrey R. Odom John

Roles Conceptualization, Data curation, Resources, Writing – review & editing

Affiliation Department of Pediatrics, The Children’s Hospital of Philadelphia, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Cynthia M. Otto

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

cmotto@vet.upenn.edu

Affiliations Penn Vet Working Dog Center, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America, Department of Clinical Sciences and Advanced Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Competing Interests

Pat Nolan owns Tactical Directional Canine which was contracted to conduct the canine training. This commercial affiliation does not alter our adherence to PLOS ONE policies on sharing data and materials.