About the Authors

Sharon S. Glaeser

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Methodology, Project administration, Validation, Writing – original draft, Writing – review & editing

Sharon.Glaeser@oregonzoo.org

Affiliation Oregon Zoo, Portland, Oregon, United States of America

Katie L. Edwards

Roles Conceptualization, Data curation, Methodology, Validation, Writing – review & editing

Affiliations Center for Species Survival, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, Front Royal, Virginia, United States of America, North of England Zoological Society, Chester Zoo, Upton-by-Chester, United Kingdom

Nadja Wielebnowski

Roles Funding acquisition, Methodology, Writing – review & editing

Affiliation Oregon Zoo, Portland, Oregon, United States of America

Janine L. Brown

Roles Conceptualization, Data curation, Funding acquisition, Methodology, Writing – review & editing

Affiliation Center for Species Survival, Smithsonian Conservation Biology Institute, Smithsonian National Zoological Park, Front Royal, Virginia, United States of America

Competing Interests

SSG is self- employed and is affiliated with the Oregon Zoo (an AZA-accredited zoo). KLE is employed by Chester Zoo, and previously by the Smithsonian Conservation Biology Institute. NW is employed by the Oregon Zoo (an AZA-accredited facility). JLB is employed by the Smithsonian Conservation Biology Institute (an AZA-accredited facility). These interests do not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.