About the Authors

Ruocheng Hu

Contributed equally to this work with: Ruocheng Hu, Yiyun Gu, Mei Luo

Roles Conceptualization, Data curation, Formal analysis, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliations Center for Nature and Society, School of Life Sciences, Peking University, Beijing, China, Shan Shui Conservation Center, Beijing, China

Yiyun Gu

Contributed equally to this work with: Ruocheng Hu, Yiyun Gu, Mei Luo

Roles Formal analysis, Methodology

Current address: Beijing Jinglang Ecology and Techniques Co., LTD., Beijing, China

Affiliation Shan Shui Conservation Center, Beijing, China

Mei Luo

Contributed equally to this work with: Ruocheng Hu, Yiyun Gu, Mei Luo

Roles Methodology, Software

Affiliation Center for Nature and Society, School of Life Sciences, Peking University, Beijing, China

Zhi Lu

Roles Conceptualization, Project administration, Supervision, Writing – review & editing

luzhi@pku.edu.cn

Affiliation Center for Nature and Society, School of Life Sciences, Peking University, Beijing, China

Ming Wei

Roles Data curation, Resources

Affiliation China Birdwatching Association, Yunnan, Kunming, China

Jia Zhong

Roles Data curation, Resources

Affiliation China Birdwatching Association, Yunnan, Kunming, China

Competing Interests

Beijing Jinglang Ecology and Techniques Co., LTD is the current employer of Ms. Yiyun Gu, but the employment relationship was established after Ms. Gu completed all her research presented in this paper. Beijing Jinglang Ecology and Techniques Co., LTD does not alter our adherence to PLOS ONE policies on sharing data and materials.