About the Authors

Matthew Traylor

Roles Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Department of Clinical Neurosciences, Stroke Research Group, University of Cambridge, Cambridge, United Kingdom, Department of Medical and Molecular Genetics, King’s College London, London, United Kingdom

Rachel Knevel

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliations Brigham and Women’s Hospital, Division of Genetics, Raychaudhuri Lab, Boston, MA, United States of America, Broad institute, Cambridge, MA, United States of America, Department of Rheumatology C1-R, Leiden University Medical Center, Albinusdreef, Leiden, the Netherlands

Jing Cui

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Division of Rheumatology Immunology and Allergy Brigham & Women's Hospital Harvard Medical School Boston, MA, United States of America

John Taylor

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Leeds Institute of Cancer & Pathology, Worsley Building Level 11 (LIDA), Clarendon Way, Leeds, United Kingdom

Westra Harm-Jan

Roles Formal analysis

Affiliations Brigham and Women’s Hospital, Division of Genetics, Raychaudhuri Lab, Boston, MA, United States of America, Broad institute, Cambridge, MA, United States of America

Philip G. Conaghan

Roles Data curation, Writing – review & editing

Affiliations Leeds Institute of Rheumatic and Musculoskeletal Medicine, University of Leeds, Leeds, United Kingdom, NIHR Leeds Biomedical Research Centre, Leeds Teaching Hospitals NHS Trust, Leeds, United Kingdom

Andrew P. Cope

Roles Conceptualization, Data curation, Funding acquisition, Writing – review & editing

Affiliation Academic Department of Rheumatology, Centre for Molecular and Cellular Biology of Inflammation, King's College London, London, United Kingdom

Charles Curtis

Roles Data curation, Writing – review & editing

Affiliations NIHR Biomedical Research Centre at South London and Maudsley NHS Foundation Trust and King’s College London, London, United Kingdom, SGDP Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, United Kingdom

Paul Emery

Roles Data curation, Writing – review & editing

Affiliations Leeds Institute of Rheumatic and Musculoskeletal Medicine, University of Leeds, Leeds, United Kingdom, NIHR Leeds Biomedical Research Centre, Leeds Teaching Hospitals NHS Trust, Leeds, United Kingdom

Stephen Newhouse

Roles Data curation, Writing – review & editing

Affiliations NIHR Biomedical Research Centre at South London and Maudsley NHS Foundation Trust and King’s College London, London, United Kingdom, Department of Biostatistics and Health Informatics, Institute of Psychiatry, Psychology and Neuroscience, King’s College London, London, United Kingdom, Farr Institute of Health Informatics Research, UCL Institute of Health Informatics, University College London, London, United Kingdom

Hamel Patel

Roles Data curation, Writing – review & editing

Affiliation NIHR Biomedical Research Centre at South London and Maudsley NHS Foundation Trust and King’s College London, London, United Kingdom

Sophia Steer

Roles Conceptualization, Data curation, Writing – review & editing

Affiliation Department of Rheumatology, King’s College Hospital, Denmark Hill, London, United Kingdom

Peter Gregersen

Roles Data curation, Writing – review & editing

Affiliation The Feinstein Institute for Medical Research, Northwell Health, Manhasset, New York, United States of America

Nancy A. Shadick

Roles Data curation, Writing – review & editing

Affiliation Division of Rheumatology Immunology and Allergy Brigham & Women's Hospital Harvard Medical School Boston, MA, United States of America

Michael E. Weinblatt

Roles Data curation, Writing – review & editing

Affiliation Division of Rheumatology Immunology and Allergy Brigham & Women's Hospital Harvard Medical School Boston, MA, United States of America

Annette Van Der Helm-van Mil

Roles Data curation, Writing – review & editing

Affiliation Department of Rheumatology C1-R, Leiden University Medical Center, Albinusdreef, Leiden, the Netherlands

Jennifer H. Barrett

Roles Conceptualization, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations NIHR Leeds Biomedical Research Centre, Leeds Teaching Hospitals NHS Trust, Leeds, United Kingdom, School of Medicine, University of Leeds, Leeds, United Kingdom

Ann W. Morgan

Roles Conceptualization, Data curation, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Leeds Institute of Rheumatic and Musculoskeletal Medicine, University of Leeds, Leeds, United Kingdom, NIHR Leeds Biomedical Research Centre, Leeds Teaching Hospitals NHS Trust, Leeds, United Kingdom

Cathryn M. Lewis

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Department of Medical and Molecular Genetics, King’s College London, London, United Kingdom, SGDP Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, United Kingdom

Ian C. Scott

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Writing – original draft, Writing – review & editing

i.scott@keele.ac.uk

Affiliations Primary Care Centre Versus Arthritis, Research Institute for Primary Care and Health Sciences, Primary Care Sciences, Keele University, Keele, United Kingdom, Haywood Academic Rheumatology Centre, Haywood Hospital, Midlands Partnership NHS Foundation Trust, High Lane, Burslem, Staffordshire, United Kingdom

Competing Interests

NAS has received less than $10,000 consultancies from Bristol myers squibb, and research grants from Mallinchrodt Pharmaceuticals and BWH (which is sponsored by Amgen, BMS, Sanofi/Regeneron, Dxterit, Crescendo and UCB). MEW has received less than $10,000 consultancies from Abbvie, Amgen, Novartis, Roche, Glaxo Smith Kline, Merck, Samsung, Crescendo Bioscience, Gilead, Pfizer and UCB, and over $10,000 from Lilly, and BMS. The BRASS study receives funding from Bristol myers squibb, Sanofi/Regeneron, AMGEN, and Crescendo Biosciences. This does not alter our adherence to PLOS ONE policies on sharing data and materials.