About the Authors

Rachel Goddard

Roles Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

rachel.goddard@jic.ac.uk (RG); paul.nicholson@jic.ac.uk (PN)

Affiliation Department of Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, England

Sarah de Vos

Roles Investigation, Writing – review & editing

Affiliation Department of Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, England

Andrew Steed

Roles Investigation, Writing – review & editing

Affiliation Department of Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, England

Amal Muhammed

Roles Investigation, Writing – review & editing

Affiliation Faculty of Applied Sciences, University of Sunderland, Sunderland, England

Keith Thomas

Roles Funding acquisition, Writing – review & editing

Affiliations Faculty of Applied Sciences, University of Sunderland, Sunderland, England, Brewlab Ltd, Sunderland Enterprise Park, Sunderland, England

David Griggs

Roles Writing – review & editing

Affiliation Crisp Malting Group Ltd, Fakenham, Norfolk, England

Christopher Ridout

Roles Conceptualization, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, England

Paul Nicholson

Roles Conceptualization, Methodology, Writing – original draft, Writing – review & editing

rachel.goddard@jic.ac.uk (RG); paul.nicholson@jic.ac.uk (PN)

Affiliation Department of Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, England

Competing Interests

Brewlab Ltd provided funding in the form of salary for author KT, and Crisp Malting Ltd provided funding in the form of salary for DG. This does not alter our adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare.