About the Authors

Manuel Plate

Roles Conceptualization, Formal analysis, Investigation, Methodology, Software, Validation, Visualization, Writing – original draft

manuel.plate@hu-berlin.de

Affiliation Institute for Bee Research Hohen Neuendorf, Hohen Neuendorf, Germany

Richard Bernstein

Roles Methodology, Software, Validation, Writing – review & editing

Affiliation Institute for Bee Research Hohen Neuendorf, Hohen Neuendorf, Germany

Andreas Hoppe

Roles Methodology, Supervision, Writing – review & editing

Affiliation Institute for Bee Research Hohen Neuendorf, Hohen Neuendorf, Germany

Kaspar Bienefeld

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – review & editing

Affiliation Institute for Bee Research Hohen Neuendorf, Hohen Neuendorf, Germany

Competing Interests

MP and AH received funding from the European Commission under its FP7 KBBE programme (2013.1.3 02, for project SmartBees Grant Agreement ‐ number 613960) (https://ec.europa.eu/research/fp7). RB received funding as part of the research program “Etablierung der genomischen Selektion zur Verbesserung von Krankheitsresistenz, Leistung, Verhalten und genetischer Vielfalt bei der Honigbiene” with project number 742 397. The project is supported by funds of the German Government’s Special Purpose Fund held at Landwirtschaftliche Rentenbank (www.rentenbank.de). The commercial affiliation does not alter our adherence to PLOS ONE policies on sharing data and material. The funders had no role in study.