About the Authors

D. Leland Taylor

Roles Formal analysis, Investigation, Writing – original draft, Writing – review & editing

Affiliations National Human Genome Research Institute, National Institutes of Health, Bethesda, United States of America, European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridgeshire, United Kingdom

David A. Knowles

Roles Investigation, Methodology, Software, Writing – review & editing

Affiliation Department of Computer Science, Stanford University, Stanford, California, United States of America

Laura J. Scott

Roles Formal analysis, Methodology, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Andrea H. Ramirez

Roles Investigation

Affiliation Department of Medicine, Vanderbilt University Medical Center, Tennessee, United States of America

Francesco Paolo Casale

Roles Methodology

Affiliation European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Brooke N. Wolford

Roles Formal analysis, Investigation

Affiliation Department of Computational Medicine & Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Li Guan

Roles Formal analysis, Investigation

Affiliation Department of Computational Medicine & Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Arushi Varshney

Roles Formal analysis, Investigation

Affiliation Department of Human Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Ricardo D’Oliveira Albanus

Roles Formal analysis, Investigation

Affiliation Department of Computational Medicine & Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Stephen C. J. Parker

Roles Investigation, Supervision

Affiliations Department of Computational Medicine & Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America, Department of Human Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Narisu Narisu

Roles Formal analysis, Investigation

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, United States of America

Peter S. Chines

Roles Formal analysis, Investigation

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, United States of America

Michael R. Erdos

Roles Investigation

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, United States of America

Ryan P. Welch

Roles Formal analysis, Investigation, Methodology

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Leena Kinnunen

Roles Investigation, Resources

Affiliation Department of Public Health Solutions, National Institute for Health and Welfare, Helsinki, Finland

Jouko Saramies

Roles Investigation, Resources

Affiliation South Karelia Social and Health Care District, Lappeenranta, Finland

Jouko Sundvall

Roles Investigation, Resources

Affiliation Department of Public Health Solutions, National Institute for Health and Welfare, Helsinki, Finland

Timo A. Lakka

Roles Investigation, Resources

Affiliations Institute of Biomedicine/Physiology, University of Eastern Finland, Kuopio, Finland, Kuopio Research Institute of Exercise Medicine, Kuopio, Finland, Department of Clinical Physiology and Nuclear Medicine, Kuopio University Hospital, University of Eastern Finland, Kuopio, Finland

Markku Laakso

Roles Investigation, Resources

Affiliations Department of Medicine, University of Eastern Finland, Kuopio, Finland, Kuopio University Hospital, Kuopio, Finland

Jaakko Tuomilehto

Roles Investigation, Resources

Affiliations Department of Public Health Solutions, National Institute for Health and Welfare, Helsinki, Finland, Department of Neurosciences and Preventive Medicine, Danube University Krems, Krems, Austria, Diabetes Research Group, King Abdulaziz University, Jeddah, Saudi Arabia, Dasman Diabetes Institute, Dasman, Kuwait

Heikki A. Koistinen

Roles Investigation, Resources

Affiliations Department of Public Health Solutions, National Institute for Health and Welfare, Helsinki, Finland, Department of Medicine and Abdominal Center: Endocrinology, University of Helsinki and Helsinki University Central Hospital, Haartmaninkatu 4, Helsinki, Finland, Minerva Foundation Institute for Medical Research, Biomedicum 2U, Tukholmankatu 8, Helsinki, Finland

Oliver Stegle

Roles Methodology, Supervision

Affiliation European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Michael Boehnke

Roles Investigation, Resources, Supervision

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Ewan Birney

Roles Investigation, Supervision, Writing – original draft, Writing – review & editing

birney@ebi.ac.uk (EB); Francis.Collins3@nih.gov (FSC)

Affiliation European Molecular Biology Laboratory, European Bioinformatics Institute, Wellcome Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Francis S. Collins

Roles Investigation, Supervision, Writing – original draft, Writing – review & editing

birney@ebi.ac.uk (EB); Francis.Collins3@nih.gov (FSC)

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, United States of America

Competing Interests

The authors have declared that no competing interests exist.