About the Authors

Sun Yajing

Contributed equally to this work with: Sun Yajing, Imran Rashid Rajput

Roles Investigation, Writing – original draft

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Imran Rashid Rajput

Contributed equally to this work with: Sun Yajing, Imran Rashid Rajput

Roles Conceptualization, Formal analysis, Writing – original draft

whliu@stu.edu.cn (LW); drimranrajput@gmail.com (IRR)

Affiliations Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China, Faculty of Veterinary and Animal Sciences, Lasbela University of Agriculture, Water and Marine Sciences, Uthal, Balochistan, Pakistan

Huang Ying

Roles Investigation

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Yu Fei

Roles Investigation

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Edmond Sanganyado

Roles Writing – review & editing

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Li Ping

Roles Software

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Wang Jingzhen

Roles Software

Affiliation Ocean College, Qinzhou University, Qinzhou, Guangxi, P.R. China

Liu Wenhua

Roles Resources, Supervision, Validation

whliu@stu.edu.cn (LW); drimranrajput@gmail.com (IRR)

Affiliation Marine Biology Institute, College of Science Shantou University, Shantou, Guangdong, P.R. China

Competing Interests

The authors have declared that no competing interests exist.