About the Authors

Pandora Pound

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Writing – original draft, Writing – review & editing


Affiliation Population Health Sciences, University of Bristol, Canynge Hall, Bristol, United Kingdom

Christine J. Nicol

Roles Investigation, Methodology, Resources, Writing – original draft, Writing – review & editing

Affiliation School of Veterinary Science, University of Bristol, Langford House, Langford, United Kingdom

Competing Interests

Pandora Pound has read the journal’s policy and the authors of this manuscript have the following competing interests: Funding from Lucy and Marius Maxwell as described above. Christine Nicol declares that no competing interests exist. This does not alter our adherence to PLOS ONE policies on sharing data and materials.