About the Authors

Shigeru Kitanishi

Roles Formal analysis, Investigation, Validation, Writing – original draft

kitanishi@oita-u.ac.jp

Current address: Faculty of Science and Technology, Oita University, Oita, Japan

Affiliation Faculty of Regional Studies, Gifu University, Gifu, Japan

Norio Onikura

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Fishery Research Laboratory, Kyushu University, Fukuoka, Japan

Takahiko Mukai

Roles Formal analysis, Funding acquisition, Investigation, Project administration, Supervision, Writing – review & editing

Affiliation Faculty of Regional Studies, Gifu University, Gifu, Japan

Competing Interests

The authors have declared that no competing interests exist.