About the Authors

G. Curtis

Contributed equally to this work with: G. Curtis, C. McGregor Argo, D. Grove-White

Roles Data curation, Formal analysis, Investigation, Methodology, Project administration, Writing – original draft, Writing – review & editing

Affiliation Department of Obesity and Endocrinology, Institute of Ageing and Chronic Disease, University of Liverpool, Leahurst Campus, Neston, Wirral, United Kingdom

C. McGregor Argo

Contributed equally to this work with: G. Curtis, C. McGregor Argo, D. Grove-White

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

Affiliation School of Veterinary Medicine, University of Surrey, Guilford, United Kingdom

D. Jones

Roles Investigation

Affiliation Institute of Veterinary Science, University of Liverpool, Leahurst Campus, Neston, Wirral, United Kingdom

D. Grove-White

Contributed equally to this work with: G. Curtis, C. McGregor Argo, D. Grove-White

Roles Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

daigw@liv.ac.uk

Affiliation Institute of Veterinary Science, University of Liverpool, Leahurst Campus, Neston, Wirral, United Kingdom

Competing Interests

The authors have declared that no competing interests exist. Whilst funding for the project was received from the following commercial organisations: Tesco Plc, Volac International and ForFarmers Plc, the authors explicitly state that these sources of funding had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript. Furthermore the authors have no commercial interests in common with any of the funders. This includes employment, consultancy, patents, products marketed or in development, or any other arrangement. This does not alter our adherence to PLOS ONE policies on sharing data and materials.