About the Authors

Clayton T. Lamb

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft

ctlamb@ualberta.ca

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Garth Mowat

Roles Data curation, Funding acquisition, Methodology, Project administration, Resources, Supervision, Validation, Writing – review & editing

Affiliations Ministry of Forests Lands and Natural Resource Operations, Nelson, British Columbia, Canada, Department of Earth, Environmental and Geographic Sciences, The University of British Columbia Okanagan Campus, Kelowna, British Columbia, Canada

Sophie L. Gilbert

Roles Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Bruce N. McLellan

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Ministry of Forests Lands and Natural Resource Operations, Nelson, British Columbia, Canada

Scott E. Nielsen

Roles Methodology, Resources, Supervision, Writing – review & editing

Affiliation Department of Renewable Resources, University of Alberta, Edmonton, Alberta, Canada

Stan Boutin

Roles Resources, Supervision, Writing – review & editing

Affiliation Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada

Competing Interests

Our project was funded by two commercial groups: BP Canada and Teck Coal, but we explicitly declare that this does not alter our adherence to PLOS ONE policies on sharing data and materials.