About the Authors

Yasunori Sugano

Roles Investigation, Validation, Writing – original draft, Writing – review & editing

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Arata Furukawa

Roles Investigation, Validation

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Osamu Nureki

Roles Validation

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Yoshiki Tanaka

Roles Validation

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Tomoya Tsukazaki

Roles Conceptualization, Investigation, Supervision, Validation, Writing – original draft, Writing – review & editing

ttsukazaki@mac.com

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Competing Interests

The authors declare the following interest: TT received funding from the Mitsubishi Foundation, a public interest incorporated foundation in Japan that is closely affiliated with the commercial entity, Mitsubishi. There are no patents, products in development, or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.