About the Authors

Yonghong Du

Roles Conceptualization, Formal analysis, Methodology, Writing – original draft

Affiliation School of Statistics, Beijing Normal University, Beijing, China

Joshua S. Martin

Roles Formal analysis, Methodology, Writing – original draft

Affiliation Department of Genetics, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

John McGee

Roles Formal analysis, Software, Writing – original draft

Affiliation NC Translational and Clinical Sciences Institute, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

Yuchen Yang

Roles Formal analysis, Software, Writing – original draft

Affiliation Department of Genetics, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

Eric Yi Liu

Roles Methodology, Software, Writing – original draft

Affiliation Department of Computer Science, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

Yingrui Sun

Roles Methodology, Writing – review & editing

Affiliation Department of Mathematics & Statistics, Boston University, Boston, Massachusetts, United States of America

Matthias Geihs

Roles Conceptualization, Formal analysis, Methodology, Software

Affiliation Department of Computer Science, Technische Universität Darmstadt, Darmstadt, Germany

Xuejun Kong

Roles Methodology, Resources, Writing – review & editing

Affiliation Beth Israel Deaconess Medical Center, Boston, Massachusetts, United States of America

Eric Lingfeng Zhou

Roles Methodology, Software, Writing – original draft

Affiliation Department of Biostatistics, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

Yun Li

Roles Conceptualization, Formal analysis, Funding acquisition, Supervision, Writing – original draft

yunli@med.unc.edu (YL); jhuang@bwh.harvard.edu (JH)

Affiliations Department of Genetics, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Computer Science, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America, Department of Biostatistics, University of North Carolina Chapel Hill, Chapel Hill, North Carolina, United States of America

Jie Huang

Roles Conceptualization, Formal analysis, Software, Supervision

yunli@med.unc.edu (YL); jhuang@bwh.harvard.edu (JH)

Affiliations Boston VA Research Institute, Boston, Massachusetts, United States of America, Brigham Women’s Hospital Division of Aging, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.