About the Authors

Lam Thanh Nguyen

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Kazunari Nakaishi

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Visualization

Affiliation TAUNS Laboratories, Inc., Izunokuni, Shizuoka, Japan

Keiko Motojima

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliation TAUNS Laboratories, Inc., Izunokuni, Shizuoka, Japan

Ayako Ohkawara

Roles Conceptualization, Data curation, Investigation, Methodology

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Erina Minato

Roles Investigation

Affiliation Laboratory of Comparative Pathology, Department of Veterinary Clinical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Junki Maruyama

Roles Investigation

Affiliation Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Takahiro Hiono

Roles Conceptualization, Investigation, Writing – review & editing

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Keita Matsuno

Roles Writing – review & editing

Affiliations Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan, Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo, Hokkaido, Japan

Masatoshi Okamatsu

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Takashi Kimura

Roles Methodology, Supervision, Writing – review & editing

Affiliation Laboratory of Comparative Pathology, Department of Veterinary Clinical Sciences, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan

Ayato Takada

Roles Supervision

Affiliation Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan

Hiroshi Kida

Roles Conceptualization, Supervision

Affiliations Research Center for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, Japan, Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo, Hokkaido, Japan

Yoshihiro Sakoda

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

sakoda@vetmed.hokudai.ac.jp

Affiliations Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Hokkaido, Japan, Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE), Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist. The TAUNS Laboratories, Inc. does not alter our adherence to PLOS ONE policies on sharing data and materials.