About the Authors

Nai-Chieh Liu

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Eileen L. Troconis

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Lajos Kalmar

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

David J. Price

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Hattie E. Wright

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Vicki J. Adams

Affiliation Vet Epi, Mildenhall, Suffolk, United Kingdom

David R. Sargan

Contributed equally to this work with: David R. Sargan, Jane F. Ladlow

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Jane F. Ladlow

Contributed equally to this work with: David R. Sargan, Jane F. Ladlow

jfl1001@cam.ac.uk

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, Cambridgeshire, United Kingdom

Competing Interests

VJA runs her own commercial consultancy company, Vet Epi. There are no patents, products in development, or marketed products to declare. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: NCL LK DRS JFL. Formal analysis: NCL LK DJP. Funding acquisition: NCL LK DRS JFL. Investigation: NCL ELT LK HEW VJA DRS JFL. Methodology: NCL LK DJP DRS JFL. Project administration: NCL LK DRS JFL. Supervision: DRS JFL. Validation: NCL ELT DRS JFL. Visualization: NCL LK DJP. Writing – original draft: NCL DRS. Writing – review & editing: NCL ELT LK DJP VJA DRS JFL.