About the Authors

Hiroaki Masuoka

Contributed equally to this work with: Hiroaki Masuoka, Kouya Shimada

Roles Data curation, Formal analysis, Investigation, Resources, Visualization, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Veterinary Public Health, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Kouya Shimada

Contributed equally to this work with: Hiroaki Masuoka, Kouya Shimada

Roles Data curation, Formal analysis, Investigation, Resources, Visualization, Writing – original draft

Affiliation Laboratory of Veterinary Public Health, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Tomoyo Kiyosue-Yasuda

Roles Conceptualization, Project administration, Resources, Supervision

Affiliation Nisshin Petfood Inc., Chiyoda-ku, Tokyo, Japan

Masaharu Kiyosue

Roles Conceptualization, Project administration, Resources, Supervision, Writing – review & editing

Affiliation Nisshin Petfood Inc., Chiyoda-ku, Tokyo, Japan

Yukie Oishi

Roles Investigation, Resources

Affiliation Nisshin Petfood Inc., Chiyoda-ku, Tokyo, Japan

Seiji Kimura

Roles Conceptualization, Funding acquisition, Project administration, Validation, Writing – review & editing

Affiliation Nisshin Petfood Inc., Chiyoda-ku, Tokyo, Japan

Yuji Ohashi

Roles Data curation, Investigation, Resources, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Food Hygiene, Department of Food and Technology, Nippon Veterinary and Life Science University, Kyonanchou, Musahino-shi, Tokyo, Japan

Tomohiko Fujisawa

Roles Data curation, Resources, Supervision, Validation, Writing – review & editing

Affiliation Laboratory of Food Hygiene, Department of Food and Technology, Nippon Veterinary and Life Science University, Kyonanchou, Musahino-shi, Tokyo, Japan

Kozue Hotta

Roles Data curation, Validation, Writing – review & editing

Affiliation Laboratory of Veterinary Public Health, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Akio Yamada

Roles Supervision, Validation, Writing – review & editing

Affiliation Laboratory of Veterinary Public Health, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Kazuhiro Hirayama

Roles Conceptualization, Data curation, Funding acquisition, Investigation, Project administration, Resources, Supervision, Validation, Writing – review & editing

akazu@mail.ecc.u-tokyo.ac.jp

Affiliation Laboratory of Veterinary Public Health, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Yayoi, Bunkyo-ku, Tokyo, Japan

Competing Interests

The present study was funded by Nisshin Petfood Inc., the employer of TKY, MK, Y. Oishi and SK. I, on behalf of all authors, state that there are no conflicts of interest exist. Nisshin Petfood Inc. only provided financial support in the form of salaries of TKY, MK, Y. Oishi and SK and research materials, but did not have additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. This does not alter our adherence to PLOS ONE policies on sharing data and materials. The specific roles of these authors are articulated in the ‘author contributions’ section.