About the Authors

Karen K. Klyczek

Contributed equally to this work with: Karen K. Klyczek, J. Alfred Bonilla, Deborah Jacobs-Sera

Roles Conceptualization, Formal analysis, Investigation, Writing – original draft, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

J. Alfred Bonilla

Contributed equally to this work with: Karen K. Klyczek, J. Alfred Bonilla, Deborah Jacobs-Sera

Roles Conceptualization, Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Deborah Jacobs-Sera

Contributed equally to this work with: Karen K. Klyczek, J. Alfred Bonilla, Deborah Jacobs-Sera

Roles Conceptualization, Formal analysis, Investigation, Writing – original draft, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Tamarah L. Adair

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Patricia Afram

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Katherine G. Allen

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Megan L. Archambault

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Rahat M. Aziz

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Filippa G. Bagnasco

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Sarah L. Ball

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Center for Life Sciences Education, The Ohio State University, Columbus, Ohio, United States of America

Natalie A. Barrett

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Robert C. Benjamin

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Christopher J. Blasi

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Katherine Borst

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Mary A. Braun

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Haley Broomell

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Conner B. Brown

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Zachary S. Brynell

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Ashley B. Bue

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Sydney O. Burke

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

William Casazza

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Julia A. Cautela

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Kevin Chen

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Nitish S. Chimalakonda

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Dylan Chudoff

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Jade A. Connor

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Trevor S. Cross

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Kyra N. Curtis

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Jessica A. Dahlke

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Bethany M. Deaton

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Sarah J. Degroote

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Danielle M. DeNigris

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Katherine C. DeRuff

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Milan Dolan

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

David Dunbar

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Marisa S. Egan

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Daniel R. Evans

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Abby K. Fahnestock

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Amal Farooq

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Garrett Finn

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Christopher R. Fratus

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Bobby L. Gaffney

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Rebecca A. Garlena

Roles Data curation, Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Kelly E. Garrigan

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Bryan C. Gibbon

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Michael A. Goedde

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Carlos A. Guerrero Bustamante

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Melinda Harrison

Roles Data curation, Supervision, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Megan C. Hartwell

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Emily L. Heckman

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biological Sciences, Lehigh University, Bethlehem, Pennsylvania, United States of America

Jennifer Huang

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Lee E. Hughes

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Kathryn M. Hyduchak

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Aswathi E. Jacob

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Machika Kaku

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Allen W. Karstens

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Margaret A. Kenna

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biological Sciences, Lehigh University, Bethlehem, Pennsylvania, United States of America

Susheel Khetarpal

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Rodney A. King

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Amanda L. Kobokovich

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Hannah Kolev

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Sai A. Konde

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Elizabeth Kriese

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Morgan E. Lamey

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Carter N. Lantz

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Jonathan S. Lapin

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Temiloluwa O. Lawson

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

In Young Lee

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Scott M. Lee

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Julia Y. Lee-Soety

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Emily M. Lehmann

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Shawn C. London

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

A. Javier Lopez

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Kelly C. Lynch

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Catherine M. Mageeney

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biological Sciences, Lehigh University, Bethlehem, Pennsylvania, United States of America

Tetyana Martynyuk

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Kevin J. Mathew

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Travis N. Mavrich

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Christopher M. McDaniel

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Hannah McDonald

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

C. Joel McManus

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Jessica E. Medrano

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Francis E. Mele

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Jennifer E. Menninger

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Sierra N. Miller

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Josephine E. Minick

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Courtney T. Nabua

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Caroline K. Napoli

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Martha Nkangabwa

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Elizabeth A. Oates

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Cassandra T. Ott

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Sarah K. Pellerino

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

William J. Pinamont

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Ross T. Pirnie

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Marie C. Pizzorno

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Emilee J. Plautz

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Welkin H. Pope

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Katelyn M. Pruett

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Gabbi Rickstrew

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Patrick A. Rimple

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Claire A. Rinehart

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Kayla M. Robinson

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Victoria A. Rose

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Daniel A. Russell

Roles Data curation, Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Amelia M. Schick

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Julia Schlossman

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Victoria M. Schneider

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Chloe A. Sells

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Jeremy W. Sieker

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Morgan P. Silva

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Marissa M. Silvi

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Stephanie E. Simon

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Amanda K. Staples

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Isabelle L. Steed

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Emily L. Stowe

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Noah A. Stueven

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Porter T. Swartz

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Emma A. Sweet

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Abigail T. Sweetman

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Corrina Tender

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Katrina Terry

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Science, Cabrini University, Radnor, Pennsylvania, United States of America

Chrystal Thomas

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Daniel S. Thomas

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Allison R. Thompson

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Lorianna Vanderveen

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Rohan Varma

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Hannah L. Vaught

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Quynh D. Vo

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Zachary T. Vonberg

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, University of Wisconsin-River Falls, River Falls, Wisconsin, United States of America

Vassie C. Ware

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biological Sciences, Lehigh University, Bethlehem, Pennsylvania, United States of America

Yasmene M. Warrad

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Kaitlyn E. Wathen

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Western Kentucky University, Bowling Green, Kentucky, United States of America

Jonathan L. Weinstein

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Saint Joseph’s University, Philadelphia, Pennsylvania, United States of America

Jacqueline F. Wyper

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biology, Baylor University, Waco, Texas, United States of America

Jakob R. Yankauskas

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Biology Department, Bucknell University, Lewisburg, Pennsylvania, United States of America

Christine Zhang

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biological Sciences, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Graham F. Hatfull

Roles Conceptualization, Formal analysis, Funding acquisition, Writing – original draft, Writing – review & editing

gfh@pitt.edu

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.