About the Authors

Ashley Case

Contributed equally to this work with: Ashley Case, Becky K. Brisson

Roles Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Becky K. Brisson

Contributed equally to this work with: Ashley Case, Becky K. Brisson

Roles Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Amy C. Durham

Roles Investigation, Methodology, Writing – review & editing

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Suzanne Rosen

Roles Investigation, Writing – original draft

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

James Monslow

Roles Investigation, Methodology

Affiliation Department of Biomedical Sciences, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Elizabeth Buza

Roles Investigation

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Pascale Salah

Roles Investigation

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Julie Gillem

Roles Investigation

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Gordon Ruthel

Roles Investigation, Methodology

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Sridhar Veluvolu

Roles Investigation

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Veronica Kristiansen

Roles Data curation, Investigation, Resources

Affiliation Department of Companion Animal Clinical Sciences, Veterinary Medicine and Biosciences, Norwegian University of Life Sciences, Oslo, Norway

Ellen Puré

Roles Investigation, Methodology, Writing – review & editing

Affiliation Department of Biomedical Sciences, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Dorothy C. Brown

Roles Formal analysis, Methodology, Software, Writing – review & editing

Affiliation Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Karin U. Sørenmo

Roles Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

swvolk@vet.upenn.edu (SWV); karins@vet.upenn.edu (KUS)

Affiliation Department of Biomedical Sciences, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Susan W. Volk

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

swvolk@vet.upenn.edu (SWV); karins@vet.upenn.edu (KUS)

Affiliations Department of Clinical Sciences and Advanced Medicine, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America, Department of Biomedical Sciences, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, PA, United States of America

Competing Interests

The authors have declared that no competing interests exist.