About the Authors

W. Bruce Saunders

wsaunder@brynmawr.edu

Affiliation Department of Geology, Bryn Mawr College, Bryn Mawr, Pennsylvania, United States of America

Emily Greenfest-Allen

Affiliation Institute for Biomedical Informatics, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Peter D. Ward

Affiliation Department of Biology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: WBS PDW. Data curation: WBS. Formal analysis: WBS EGA. Investigation: WBS PDW. Resources: WBS PDW. Visualization: WBS EGA. Writing – original draft: WBS. Writing – review & editing: WBS EGA.