About the Authors

Jieming Shi

Contributed equally to this work with: Jieming Shi, Xi Li

Affiliation Department of Biology, Miami University, Oxford, Ohio, United States of America

Xi Li

Contributed equally to this work with: Jieming Shi, Xi Li

Affiliations Department of Biology, Miami University, Oxford, Ohio, United States of America, College of Information Science and Engineering, Guangxi University for Nationalities, Nanning, Guangxi, China

Min Dong

Affiliations Department of Biology, Miami University, Oxford, Ohio, United States of America, Department of Automation, Xiamen University, Fujian, China

Mitchell Graham

Affiliation Department of Biology, Miami University, Oxford, Ohio, United States of America

Nehul Yadav

Affiliation Department of Biology, Miami University, Oxford, Ohio, United States of America

Chun Liang

liangc@miamioh.edu

Affiliation Department of Biology, Miami University, Oxford, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: CL. Formal analysis: JS. Funding acquisition: CL. Investigation: CL. Methodology: JS XL CL. Project administration: CL. Software: JS XL MD MG NY. Supervision: CL. Writing – original draft: JS XL CL. Writing – review & editing: JS XL CL.