About the Authors

Marion Desclozeaux

Affiliation Centre for Animal Health Innovation, Faculty of Science, Health, Education and Engineering, University of the Sunshine coast, Maroochydore DC, Queensland, Australia

Amy Robbins

Affiliation Endeavour Veterinary Ecology Pty Ltd, Toorbul, Queensland, Australia

Martina Jelocnik

Affiliation Centre for Animal Health Innovation, Faculty of Science, Health, Education and Engineering, University of the Sunshine coast, Maroochydore DC, Queensland, Australia

Shahneaz Ali Khan

Affiliation Centre for Animal Health Innovation, Faculty of Science, Health, Education and Engineering, University of the Sunshine coast, Maroochydore DC, Queensland, Australia

Jon Hanger

Affiliation Endeavour Veterinary Ecology Pty Ltd, Toorbul, Queensland, Australia

Volker Gerdts

Affiliation Vaccine and Infectious Disease Organization– International Vaccine Centre, University of Saskatchewan, Saskatoon, Saskatchewan, Canada

Andrew Potter

Affiliation Vaccine and Infectious Disease Organization– International Vaccine Centre, University of Saskatchewan, Saskatoon, Saskatchewan, Canada

Adam Polkinghorne

Affiliation Centre for Animal Health Innovation, Faculty of Science, Health, Education and Engineering, University of the Sunshine coast, Maroochydore DC, Queensland, Australia

Peter Timms

ptimms@usc.edu.au

Affiliation Centre for Animal Health Innovation, Faculty of Science, Health, Education and Engineering, University of the Sunshine coast, Maroochydore DC, Queensland, Australia

Competing Interests

AR and JH were employed by Endeavour Veterinary Ecology. This does not alter our adherence to PLOS ONE policies on sharing data and materials. PT and AP received funding from the Australian Research Council (ARC) for this project, however the ARC did not play a role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript and only provided financial support in the form of authors' salary (MD) and research materials.

Author Contributions

Conceptualization: PT A. Polkinghorne VG A. Potter MD. Data curation: MD AR MJ PT A. Polkinghorne. Formal analysis: MD MJ PT A. Polkinghorne. Funding acquisition: PT A. Polkinghorne JH. Investigation: MD AR MJ SAK PT A. Polkinghorne. Methodology: MD MJ PT. Project administration: PT MD JH. Resources: PT A. Polkinghorne JH VG A. Potter. Supervision: PT A. Polkinghorne. Validation: MD MJ SAK JH. Writing – original draft: MD JH A. Polkinghorne PT. Writing – review & editing: MD PT A. Polkinghorne MJ JH VG A. Potter SAK AR.